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Late Hargobind Khorana had received Nobel Prize in physiology in 1968, for work on which among the following?

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Late Hargobind Khorana had received Nobel Prize in physiology in 1968, for work on which among the following?

1) Mechanisms of the action of hormones
2) Chemical Structure of antibodies
3) Interpretation of the genetic code
4) Structural and functional organization of the cell

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